There is one main tank that uses transfer pumps to feed fuel into two supply tanks. The supply tanks in turn feed the engines (one tank per engine).
The supply tanks were found empty, whilst 76Kg fuel remained in the main tank.
During flight the volume of fuel in the main tank naturally reduces. Once the fuel reduces below a certain level it leads to situations where a transfer pump can be "out of the water" i.e in the hover the nose is up, fuel in the main tank sloshes rearwards and the fwd pump is uncovered. The instruction in the manual is to turn the fwd transfer pump OFF whilst in this situation to avoid damaging the pump. Once the cab transitions into forward flight the attitude is nose down meaning the fuel sloshes to the front of the tank. In this case the manual says that the fwd transfer pump should be turned back ON (as it is now immersed in fuel) and the rear transfer pump can be turned OFF (as it is now above the fuel level).
Due to the nature of the mission they would have spent time travelling to a tasking (nose down) then hovering at a tasking (nose up), repeat X many times. As the mission continued and fuel in the main tank reduced, the pilot would have to switch the fwd and rear transfer pumps ON/OFF several times. Eventually, both transfer pumps ended up OFF, meaning no fuel from the main tank reaching the supply tanks. Once fuel in the supply tanks run out, engines flame.
But there should have been lots of warnings before this happened.